Published March 2018. Copyright 2018 by Van Andel Institute: all rights reserved. Van Andel Institute, 333 Bostwick Avenue, N.E. Grand Rapids, Michigan 49503, U.S.A. ii | VAN ANDEL RESEARCH INSTITUTE SCIENTIFIC REPORT
In Memoriam Arthur S. Alberts, Ph.D. 1964–2016 Art Alberts passed away in December 2016 after a courageous eight-year battle with brain cancer. He was a passionate, deeply inquisitive scientist who joined VARI in 2000 as one of its first scientific investigators. Art was brought up in Southern California, but he never seemed to allow Michigan winters to intimidate him into forgoing flip-flops and shorts. He was a friend, mentor, and collaborator, a man who loved the purity of science, the thrills of a dangerous mountain bike trail, and a good IPA. He is deeply missed. VAN ANDEL RESEARCH INSTITUTE SCIENTIFIC REPORT | iii
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